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Exercise

workout
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Mathematical modelling in biology, Assignment 1

Dr. Aziz Ouhinou, AIMS South Africa

March 3, 2014

1 Harvesting a single natural population


Consider a model which consists of a harvest strategy that depends linearly on the total population N (t)
with a constant harvesting rate e:
dN N
= rN (1 − ) − eN = f (N ).
dt K
The parameters r, K and e are positive.

1. Find the steady states (discuss with respect to parameters) and analyse their local stability.

2. Present the stability diagram. Discuss the obtained results.

3. Give the maximum sustainable yield (Y = eN ). Conclude.

2 Control of insect outbreaks


An insect population satisfies the equation

dv v2
= − βv
dt 1 + v2
where β is a positive parameter. Determine the steady states and their linear stability. Show that when
β is very small, v has a very large stable steady state. Sketch a graph of how this steady state value will
vary as β increases.

3 Demography of baleen whales


A continuous time model for the baleen whale (a slightly more complicated model of which the interna-
tional whaling commission used) is the delay equation

dN
= −µN (t) + µN (t − T )[1 + q(1 − (N (t − T )/K)z )].
dt
Here µ(> 0) is a measure of the mortality, q(> 0) is the maximum increase in fecundity the population
is capable of, K is the unharvested carrying capacity, T is the time to sexual maturity and z > 0 is a
measure of the intensity of the density-dependent response as the population drops.

1
1. Determine the steady state populations. Show that the linearised equation around the positive
steady state is
dn(t)
= −µn(t) − µ(qz − 1)n(t − T ).
dt
2. Find that the positive steady state is locally stable if
1
π − arccos( )
µT < µTc = p 2 b , b = qz − 1 > 1
(b − 1)

and stable for all T if b < 1.

3. For b > 1 and Tc < T < Tc + , where (  1), show that the period of oscillations is
proximately equal √2π .
µ b2 −1

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