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After the events of Psycho-Pass 3, Inspector Kei Mikhail Ignatov finds himself involved with an organization named Bifrost. First Inspector Koichi Azusawa coordinates an assault on the Public Safety Bureau tower.

2112; the summer before Akane Tsunemori was assigned to Criminal Investigation Department. Teppei Sugo, an accomplished pilot of the Defense Army, joins the military operation in Okinawa. Three months later, an unmanned combat drone opens fire on the Ministry of Defense in Tokyo. Enforcer Tomomi Masaoka is dispatched to Sugo's military base to investigate the truth behind this case.